Three trees of 21 skills each (63 in all): Replication, Transmission and Persistence. Each has three columns, its own resource, an active skill with augments, and three capstones to choose from.
Resource: Reservoir. 40% of the damage you take is stored. Kills and room clears pour it back as healing.
Active skill: Seasonal Rebound
Capstones (pick 1): Fortified Provirus, Every Winter, Undying Strain
From the Codex
The protein shell that encloses a virus's genome, built from repeating subunits called capsomeres. Most are icosahedral (20 triangular faces) or helical.
Fun fact: Capsids self-assemble: the proteins snap together on their own, like a puzzle that builds itself.
Each tree has 6 rows and a capstone row. A row opens once you've spent enough points in that tree, and then every skill in it is open: no skill needs another one first.
Rows open at 0, 5, 8, 11, 14 and 17 points spent in that tree; the capstone row at 21
Every rank costs 1 skill point
Rows also need a virus level, finished generations, and for the deep rows the furthest organ reached (organ 2, 4, 6 and 8)
From the Codex
Your virus's experience and the skills it buys.
Fun fact: Switching to another capstone in the same tree is free: its point moves over.
Resources, kill streaks and when-you-get-hit skills show as round icons under your health: a small number for the stacks and a clock hand that darkens the icon as its time runs out.
When-you-get-hit skills (Pressure Response, Startle Response, Inflammation) stack 10 times and last 20 s, each hit restarting the timer
From the Codex
Your virus's experience and the skills it buys.
Fun fact: Switching to another capstone in the same tree is free: its point moves over.