
Max HP
Capsid · Envelope · MatrixShown as '+N% max HP'.
- About +6% on Common, about +20% on Prismatic
From the Codex
Stabilizing mutations in capsid proteins help virions survive heat, drying and stomach acid.
Robust Capsid in the Codex
Rolls
A piece rolls as many stats as its tier allows (1 on Common, 7 on Prismatic, out of the 7 or 8 its slot can roll). Each value lands within 20% of a base, times the tier's roll strength, then grows with enhancing, item level and mutations. The same stat on several pieces adds up, then applies once. Typical values below are before enhancing.

Shown as '+N% max HP'.
Stabilizing mutations in capsid proteins help virions survive heat, drying and stomach acid.
Robust Capsid in the Codex
Shown as '+N% damage'.
Your virus fires on its own at the nearest cell it can see.
Fun fact: Positioning is your aim: stand where the cell you want is the nearest one in sight.
Auto-Attack & Shot Types in the Codex
Shown as '+N% move speed'.
Some viruses shed non-essential genes, letting them copy themselves faster and spread sooner.
Streamlined in the Codex
Shown as '+N% fire rate'.
A faster polymerase means more copies of the genome per hour.
Rapid Replication in the Codex
Shown as '+N% ultimate charge'.
Your three special buttons on the right.
Fun fact: Save the ultimate for bosses and big waves.
Signature, Ultimate & Skill in the Codex
Charges your signature ability faster. Shown as '+N% Shift charge', or 'Latency charge' for Retrovirus.
An abrupt, major change in a flu virus, caused when two strains infect the same cell and swap RNA segments (reassortment).
Fun fact: Shifts have produced new pandemic strains, including those of 1957, 1968 and 2009.
Antigenic Shift in the Codex
Shown as '−N% damage taken'.
Extra health bars that soak damage before a cell's health.
Fun fact: Match your damage to the defence: electric for shields, explosive for armor.
Shields, Armor & Damage Types in the Codex
Shown as 'Auto-attacks heal N% of their damage'.
Lucky double-damage hits, and healing from your own damage.
Fun fact: Lifesteal only works on auto-attacks, so it rewards keeping your fire up.
Critical Hits & Lifesteal in the CodexShown as 'Regenerate N HP/s'.

Shown as '+N% pickup range'.
High-affinity attachment proteins help virions stick to and gather around target cells.
Magnetic Capsid in the Codex
Extra damage on hits against cells that haven't noticed you yet. Shown as '+N% ambush damage'.
Hitting a cell before it has noticed you.
Fun fact: Gear, strains and skill nodes can push ambush damage even higher.
Ambush in the Codex
Shown as 'Detection rises N% slower'.
Many viruses spread before symptoms appear, staying under the immune system's radar.
Silent Spread in the Codex
Shown as '+N% critical hit chance'.
Lucky double-damage hits, and healing from your own damage.
Fun fact: Lifesteal only works on auto-attacks, so it rewards keeping your fire up.
Critical Hits & Lifesteal in the Codex
Shown as '+N% critical hit damage'.
Lucky double-damage hits, and healing from your own damage.
Fun fact: Lifesteal only works on auto-attacks, so it rewards keeping your fire up.
Critical Hits & Lifesteal in the Codex