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Skill tree · Tree 1

Replication Tree

Offence. Copy faster than the immune system can respond: build Viral Load by hitting cells, then cash it in. Resource: Viral Load. Active skill: Viral Burst.

On this page18
  1. Point Mutation
  2. Viral Burst
  3. Viral Load
  4. Epitope Shift
  5. Double Capsid
  6. Replication Frenzy
  7. Antibody Escape
  8. Shrapnel Ring
  9. Load Bearing
  10. Marked Target
  11. Cytokine Echo
  12. Pressure Response
  13. Escape Variant
  14. Twin Strain
  15. Hypermutation
  16. Lethal Mutation
  17. Burst Memory
  18. Error Catastrophe

Point Mutation

Row 1 · Stat · Copy Errors

+4% damage (per rank)

  • 5 ranks, 1 point each (max: +20% damage)
  • Opens at 0 points in the tree, level 1
From the Codex

Your virus's experience and the skills it buys.

Fun fact: Switching to another capstone in the same tree is free: its point moves over.

Levels & Skill Tree in the Codex

Viral Burst

Row 1 · Active skill · Viral Burst

Fire a ring of 12 shots around you, each at full damage · 9s cooldown

  • 1 rank, 1 point
  • Opens at 0 points in the tree, level 1
From the Codex

Your virus's experience and the skills it buys.

Fun fact: Switching to another capstone in the same tree is free: its point moves over.

Levels & Skill Tree in the Codex

Viral Load

Row 1 · Resource · Overload

Unlocks Viral Load: each hit adds a stack, up to 10, and they drain after 3s without a hit. +1% fire rate per stack

  • 1 rank, 1 point
  • Opens at 0 points in the tree, level 1
From the Codex

Your virus's experience and the skills it buys.

Fun fact: Switching to another capstone in the same tree is free: its point moves over.

Levels & Skill Tree in the Codex

Epitope Shift

Row 2 · Stat · Copy Errors

+3% critical hit chance (per rank)

  • 5 ranks, 1 point each (max: +15% critical hit chance)
  • Opens at 5 points in the tree, level 3
From the Codex

Your virus's experience and the skills it buys.

Fun fact: Switching to another capstone in the same tree is free: its point moves over.

Levels & Skill Tree in the Codex

Double Capsid

Row 2 · Augment · Viral Burst

Viral Burst fires a second ring 0.3s after the first, offset so the gaps are covered

  • 1 rank, 1 point
  • Opens at 5 points in the tree, level 3
From the Codex

Your virus's experience and the skills it buys.

Fun fact: Switching to another capstone in the same tree is free: its point moves over.

Levels & Skill Tree in the Codex

Replication Frenzy

Row 2 · Kill skill · Overload

Each kill gives +6% fire rate for 4s. Stacks up to 3 times (per rank)

  • 3 ranks, 1 point each (max: Each kill gives +18% fire rate for 4s. Stacks up to 3 times)
  • Opens at 5 points in the tree, level 3
From the Codex

Your virus's experience and the skills it buys.

Fun fact: Switching to another capstone in the same tree is free: its point moves over.

Levels & Skill Tree in the Codex

Antibody Escape

Row 3 · Conditional · Copy Errors

+12% damage to cells you haven't hit yet (per rank)

  • 3 ranks, 1 point each (max: +36% damage to cells you haven't hit yet)
  • Opens at 8 points in the tree, level 6, 1 finished generation
From the Codex

Your virus's experience and the skills it buys.

Fun fact: Switching to another capstone in the same tree is free: its point moves over.

Levels & Skill Tree in the Codex

Shrapnel Ring

Row 3 · Augment · Viral Burst

Viral Burst shots pierce 2 cells and split in two on their last hit

  • 1 rank, 1 point
  • Opens at 8 points in the tree, level 6, 1 finished generation
From the Codex

Your virus's experience and the skills it buys.

Fun fact: Switching to another capstone in the same tree is free: its point moves over.

Levels & Skill Tree in the Codex

Load Bearing

Row 3 · Resource · Overload

Viral Load also gives +0.8% damage per stack (per rank)

  • 3 ranks, 1 point each (max: Viral Load also gives +2.4% damage per stack)
  • Opens at 8 points in the tree, level 6, 1 finished generation
From the Codex

Your virus's experience and the skills it buys.

Fun fact: Switching to another capstone in the same tree is free: its point moves over.

Levels & Skill Tree in the Codex

Marked Target

Row 4 · Conditional · Copy Errors

Critical hits mark a cell for 3s. Marked cells take +6% damage from everything (per rank)

  • 3 ranks, 1 point each (max: Critical hits mark a cell for 3s. Marked cells take +18% damage from everything)
  • Opens at 11 points in the tree, level 10, 3 finished generations, reach organ 2 (Lungs)
From the Codex

Your virus's experience and the skills it buys.

Fun fact: Switching to another capstone in the same tree is free: its point moves over.

Levels & Skill Tree in the Codex

Cytokine Echo

Row 4 · Augment · Viral Burst

Each cell Viral Burst kills refunds 0.8s of its cooldown, up to half of it per cast (per rank)

  • 3 ranks, 1 point each (max: Each cell Viral Burst kills refunds 2.4s of its cooldown, up to half of it per cast)
  • Opens at 11 points in the tree, level 10, 3 finished generations, reach organ 2 (Lungs)
From the Codex

Your virus's experience and the skills it buys.

Fun fact: Switching to another capstone in the same tree is free: its point moves over.

Levels & Skill Tree in the Codex

Pressure Response

Row 4 · Conditional · Overload

Each time you get hit: +2% projectile speed. Stacks 10 times; the stacks last 20s (each hit restarts the timer) (per rank)

  • 3 ranks, 1 point each (max: Each time you get hit: +6% projectile speed. Stacks 10 times; the stacks last 20s (each hit restarts the timer))
  • Opens at 11 points in the tree, level 10, 3 finished generations, reach organ 2 (Lungs)
From the Codex

Your virus's experience and the skills it buys.

Fun fact: Switching to another capstone in the same tree is free: its point moves over.

Levels & Skill Tree in the Codex

Escape Variant

Row 5 · Conditional · Copy Errors

+6% damage to cells under 35% health (per rank)

  • 5 ranks, 1 point each (max: +30% damage to cells under 35% health)
  • Opens at 14 points in the tree, level 15, 6 finished generations, reach organ 4 (Lymph Node)
From the Codex

Your virus's experience and the skills it buys.

Fun fact: Switching to another capstone in the same tree is free: its point moves over.

Levels & Skill Tree in the Codex

Twin Strain

Row 5 · Augment · Viral Burst

Tap the skill button for the ring as usual, or hold it to charge a cone blast in your aim direction (up to 2x damage)

  • 1 rank, 1 point
  • Opens at 14 points in the tree, level 15, 6 finished generations, reach organ 4 (Lymph Node)
From the Codex

Your virus's experience and the skills it buys.

Fun fact: Switching to another capstone in the same tree is free: its point moves over.

Levels & Skill Tree in the Codex

Hypermutation

Row 5 · Resource · Overload

At full Viral Load, every 4th shot is a guaranteed critical hit

  • 1 rank, 1 point
  • Opens at 14 points in the tree, level 15, 6 finished generations, reach organ 4 (Lymph Node)
From the Codex

Your virus's experience and the skills it buys.

Fun fact: Switching to another capstone in the same tree is free: its point moves over.

Levels & Skill Tree in the Codex

Lethal Mutation

Row 6 · Stat · Copy Errors

+8% critical hit damage (per rank)

  • 5 ranks, 1 point each (max: +40% critical hit damage)
  • Opens at 17 points in the tree, level 19, 8 finished generations, reach organ 6 (Liver)
From the Codex

Your virus's experience and the skills it buys.

Fun fact: Switching to another capstone in the same tree is free: its point moves over.

Levels & Skill Tree in the Codex

Burst Memory

Row 6 · Stat · Viral Burst

Viral Burst cooldown -6% (per rank)

  • 5 ranks, 1 point each (max: Viral Burst cooldown -30%)
  • Opens at 17 points in the tree, level 19, 8 finished generations, reach organ 6 (Liver)
From the Codex

Your virus's experience and the skills it buys.

Fun fact: Switching to another capstone in the same tree is free: its point moves over.

Levels & Skill Tree in the Codex

Error Catastrophe

Row 6 · Risk · Overload

+25% damage, but you take 15% more damage

  • 1 rank, 1 point
  • Opens at 17 points in the tree, level 19, 8 finished generations, reach organ 6 (Liver)
From the Codex

Your virus's experience and the skills it buys.

Fun fact: Switching to another capstone in the same tree is free: its point moves over.

Levels & Skill Tree in the Codex

See also